WEBVTT

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You know, there is something deeply permanent

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about a name. Oh, for sure. You walk into a museum,

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you see a massive skeleton mounted on a frame.

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Yeah. And there's a little brass plaque. It says

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Tyrannosaurus Rex or Triceratops. And you just,

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you trust that plaque. Right. You assume it's

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settled science. Exactly. You assume that name

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is as solid as the rock the bone was dug out

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of. But what if I told you that in paleontology,

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names are sometimes, let's say, written in pencil?

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Or in the case we are looking at today, written

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in air quotes. Yes, that is the thing that grabbed

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me immediately when I opened up the packet for

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this deep dive. It stands out, doesn't it? It

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really does. Because we're looking at a creature

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from about 304 million years ago, but the title

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of the research isn't just Haptodus garnetensis.

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It is Haptodus, with these big, aggressive quotation

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marks around it, garnetensis. It honestly looks

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like the scientific record is being sarcastic.

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It totally does look that way. But those quotation

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marks are actually doing a lot of heavy lifting.

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How so? They are basically a signal flare. They

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tell us that we are stepping into this crime

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scene of mistaken identity, bureaucratic red

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tape, and a decades -long argument about who

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our ancestors actually were. So it is our mission

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for this deep dive. We're going to unpack the

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mystery of the quotation marks. We're going to

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travel back to ancient Kansas to look at a fossil

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that has been having a literal identity crisis

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since the 1970s. A very long identity crisis.

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Right. And spoiler alert for you listening. It

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turns out Haptotus might not be one animal. It

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might be three different animals wearing the

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same trench coat. A very dusty 300 million year

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old trench coat. Exactly. So let's set the stage

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before we get into all the Latin names and the

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taxonomy wars. We are not in the Jurassic Park

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era here. We are way further back. Much further.

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We are in the late Carboniferous period, specifically

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the late Cosimovian stage. So we are talking

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roughly 304 million years ago. And geographically,

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we are in Anderson County, Kansas. The site is

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called the Garnett Quarry. Now, I've driven through

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Kansas. I picture flat horizons, wheat fields,

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maybe a storm cellar. Sure, the classic Midwest.

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Yeah. But if I step out of the time machine in

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the Casamovian stage, is that what I'm seeing?

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Absolutely not. You are looking at a completely

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different planet, essentially. Okay. In the late

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Carboniferous, Kansas was sitting very close

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to the equator. It would have been hot, likely

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very humid, though the climate was just starting

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to fluctuate a bit. The Garnett Quarry site specifically

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represents a coastal or lagoonal environment.

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So water. Water and mud. Lots and lots of mud.

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A prehistoric spa day. If the spa was full of

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primitive conifers and teeming with early tetrapods,

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sure. It was this thriving terrestrial ecosystem.

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And that mud is super important because it acts

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like a time capsule. Because things get stuck

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in it. Right. It trapped these animals and preserved

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them in incredible detail within what geologists

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call the rock lake member of the Stanton Formation.

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Okay, so we are standing in the equatorial mud

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of ancient Kansas. We see a creature scuttling

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by. Let's look at the so -called Haptotis garnetensis.

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If I see this thing, what is my first instinct?

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Do I scream lizard? Do I scream dinosaur? If

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you scream dinosaur, you'd be about 70 or 80

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million years too early. Fair enough. And if

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you screamed lizard, you'd be describing its

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general vibe. I mean, it sprawls. It has a long

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tail. It looks reptilian. But you would be dead

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wrong about its family tree. Because this animal

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is a synapsid. Yes, exactly. It is a synapsid.

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Let's unpack synapsid for a second. We hear this

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term, but I think people usually just gloss over

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it as weird reptile thing. Which is a shame because

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it's probably the most important distinction

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in vertebrate history. Imagine a fork in the

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road. Life crawls out of the water. Eventually,

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there's a massive evolutionary split. Okay. One

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road leads to the cladesaur opsida. That path

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gives you reptiles, lizards, dinosaurs, and eventually

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birds. That's the path we usually see in the

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movies, the Jurassic Park path. Right. But there

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is another road. The synapsida. That is the line

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that leads to thrapsids, cynodonts, and eventually

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mammals. Us. Us. So when you look at our little

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Haptotis garnetensis, you aren't looking at a

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lizard's great -grandfather. You are looking

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at a very, very distant great -uncle on your

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own family tree. We call them stem mammals, right?

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They do. I love that concept. It's team mammal

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in a disguise. It looks like a reptile. Acts

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like a reptile. But deep down, its biology is

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pointing toward humanity. That is a great way

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to put it. Specifically, this animal is what

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we call a basal sphenacodont. Sphenacodont. I

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feel like I know that word from somewhere. You

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probably do, mostly thanks to toy stores. The

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most famous sphenacodont is Dimetrodon. Oh. The

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one with the giant sail on its back? That's the

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one. The one that is always included in the bag

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of plastic dinosaurs, even though... It definitely

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isn't a dinosaur. We're very same. Dimetrodon

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is kind of the rock star of the sphenacodons.

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Now, Argi, the haptotis from Kansas, is related

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to Dimetrodon, but it is basal. Meaning lower

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on the tree. Right. It sits lower on the evolutionary

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tree. It's an earlier branch, so it didn't have

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the big flashy sail. It was much more modest.

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So it's like the beta version. Before they added

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the spoiler and the raising stripes. In a way,

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yes. But it was a highly successful design. These

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were predators. They had sharp teeth, strong

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jaws. They were eating insects, maybe small amphibians.

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They were doing quite well in that Kansas lagoon.

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And the reason we know they were doing well is

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because we didn't just find one random toe bone.

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The source material for this deep dive makes

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a massive deal about the quality of the fossils

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found at Garnet Quarry. It really is a paleontologist's

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dream. Usually from this specific era, you get

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fragments. A jaw piece here, a crushed rib there.

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But at Garnet, they found an absolute treasure

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trove. The source highlights a specific holotype

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specimen, RM14156. Now, a holotype is the standard

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for this species, right? Like the reference model.

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Correct. It's the physical ruler you measure

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every other fob against. This specimen, RM14156,

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was a partially articulated skeleton. It had

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both cranial parts, so the skull and post -cranial

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parts, the body, all together. Which is rare

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enough. Very rare. But they didn't stop there.

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Over time, they found more than 16 well -preserved

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specimens in that quarry. And here. is where

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it gets really interesting to me. The notes mention

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that the holotype, that gold standard specimen,

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was a teenager. It was an immature individual,

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yeah. And that is actually a huge stroke of luck

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for us. Because we have so many specimens from

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the site, we have what we call an ontogenetic

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series. Ontogenetic. That sounds like an expensive

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word. It basically just means growing up. We

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have fossils that represent different stages

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of the animal's life. We can see the babies,

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the adolescents, and the full -grown adults.

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That must be incredibly rare. It is. Usually

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you find one bone from one adult and you have

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to guess what the rest of the life cycle looked

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like. flip through the family photo album so

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we have a prehistoric nursery we have incredible

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fossils we know where they lived we know there

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are distant mammalian cousins so why the identity

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crisis why the scare quotes in the name this

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is where we leave the mud of Kansas and enter

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the dusty libraries of taxonomy this is the taxonomy

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war I assume this war started when the creature

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was first named It started with perfectly good

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intentions. Back in 1977, a paleontologist named

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Currie described these fossils. At the time,

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looking at the general anatomy, they seemed to

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fit neatly into an already existing genus called

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Haptotus. Right. So he named the Kansas animals

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Haptotus garnetensis, which just means the Haptotus

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from Garnet. 1977. Simple times. Everyone is

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happy. For a while, yeah. But science is a process

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of refinement. Fast forward to the early 90s,

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1993 and 1994 to be exact, a researcher named

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Michel Loren decided to take a much closer look.

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But he didn't just look at the Kansas fossils.

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He looked at the foundation of the Heptotis genus

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itself. And he found cracks in the foundation.

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Massive ones. Every genus has a type species.

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That's the original definition. For Heptotis,

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the type species was Heptotis balei, which was

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found in Europe. Lauren went back to those original

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bailey fossils and realized they were, well,

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to put it politely, they were insufficient. Insufficient

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how? They were just too fragmentary. They didn't

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have enough unique anatomical features to really

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define a group. So Lauren declared Haptotis bailey

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a nomandubium. A nomandubium. Yeah. A dubious

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name. Exactly. A doubtful name. Now think about

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the logical cascade here. If the definition of

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haptotis is based on a pile of broken rocks from

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Europe that can't actually be identified, then

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the name itself is hollow. It's a phantom. It

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is. And if the genus Heptodus is a phantom, you

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can't attach our nice, solid, beautifully preserved

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Kansas fossils to it. So Heptodus garnetensis

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is essentially evicted. It's wearing a name tag

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that belongs to a ghost. Precisely. And that

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is exactly why the quotation marks appeared.

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The scientific community was saying, look, we

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know this isn't really a Heptodus because Heptodus

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barely exists anymore, but we don't have a new

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name for the Kansas fossils yet. So air quotes.

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That seems incredibly frustrating. It's like

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living in a... house for 20 years, but never

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being on the lease. It is frustrating, but it

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protects the scientific record from chaos. You

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wait until someone does the rigorous work to

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fix it. And that brings us to 2015 and a researcher

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named Friedrich Spindler. Spindler is kind of

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the hero of our story today. He decides to finally

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clean up this mess. He does. For his dissertation,

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Spindler did a massive deprovision of these basal

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synapsids. He compared the Kansas fossils to

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the European types and confirmed what Lauren

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had suspected all along. That they weren't the

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same. Right. The Kansas animal was definitely

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not congeneric with the original Haptodus. It

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was completely distinct. It needed its own name.

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And he gave it one. He proposed Eohaptodus. Eohaptodus,

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which translates roughly to Dawn Haptodus. Okay,

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so problem solved. We have a new name. We can

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take down the quotation marks. Why does the source

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material still use them? Why aren't we doing

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a deep dive on Eohaptodus instead of so -called

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Haptodus? Because of the bureaucracy of science.

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Of course. Spindler proposed the name Eohaptodus

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in his dissertation. But under the International

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Code of Zoological Nomenclature, which is basically

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the strict rulebook for naming animals, a dissertation

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is not always considered a formal publication.

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You're kidding. So because it was in a thesis

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and not, say, the Journal of Vertebrate Paleontology,

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it doesn't count? It counts as a very strong

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proposal, but not as an official law. So the

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name Eohaptidus is currently considered a nomen

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ex dissertatione. Nomen ex dissertatione. Literally

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a name from a dissertation. Exactly. Think of

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it as a waiting room name. It's valid in the

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sense that scientists know exactly what you're

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talking about when you say it, but it hasn't

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received that final formal stamp of approval

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to become official. So the quotation marks stay

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up as a warning sign. Like, caution, this name

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is pending validation. You got it. I actually

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kind of love that. It shows that science isn't

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just about discovery. It's about peer review,

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consensus, and making sure the filing system

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works. It's slow for a reason. It definitely

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is. But here is where the story gets really wild.

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Spindler didn't just stop at renaming the main

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group. He kept looking. Right. And this is the

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plot twist. Because up until this point... For

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40 years, everyone assumed that all 16 specimens

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in that Kansas quarry were the exact same animal.

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Just a big happy family of Haptidis or Eohaptidis

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hanging out in the mud together. That was the

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baseline assumption. One site, one species. But

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in 2020, Spindler went back to the bones. He

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started looking incredibly closely at specific

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specimens that had just been lumped in with the

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others for decades. And he noticed differences

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that couldn't be explained away by saying, oh,

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it's just a teenager or, oh, it's a female. These

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were fundamental anatomical differences. Correct.

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He looked at a specific specimen labeled ROM43608.

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Previously, everyone just nodded and said, yep,

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another Heptotis. But Spindler looked at the

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jaw structure, he looked at the teeth, and realized,

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this is a completely different animal. So there

00:12:14.149 --> 00:12:16.169
was a stranger in the nursery. A total stranger.

00:12:16.190 --> 00:12:18.690
He realized it was a new genus entirely, and

00:12:18.690 --> 00:12:21.350
he named it Kinomagnathus. Kinomagnathus. It's

00:12:21.350 --> 00:12:23.549
a robust name for a very robust animal. Yeah.

00:12:23.590 --> 00:12:25.529
But he wasn't done there. He looked at another

00:12:25.529 --> 00:12:30.090
specimen, ROM 43601. Let me guess. Not a Haptotis

00:12:30.090 --> 00:12:32.950
either. Not a Haptotis. He found distinct features

00:12:32.950 --> 00:12:35.210
in the skull and the limbs that separated it

00:12:35.210 --> 00:12:37.610
from both the main group and from Ceno -Magnathus.

00:12:37.750 --> 00:12:41.450
He split this one out as Tenuocaptor razi. Tenuocaptor.

00:12:41.970 --> 00:12:45.409
And just like Eohaptotis, the notes mention that

00:12:45.409 --> 00:12:48.389
this name is also a Nomen ex dissertationae,

00:12:48.490 --> 00:12:51.370
so it's also sitting in the lobby waiting for

00:12:51.370 --> 00:12:54.870
its official badge. It is. But don't let the

00:12:54.870 --> 00:12:57.509
legal bureaucratic status distract you from the

00:12:57.509 --> 00:13:00.269
biological bombshell here. We went from thinking

00:13:00.269 --> 00:13:03.029
this specific quarry contained one single species

00:13:03.029 --> 00:13:05.629
to realizing it contained three entirely different

00:13:05.629 --> 00:13:08.789
animals. That is wild. You had Eohaptidis, the

00:13:08.789 --> 00:13:11.610
main group. You had Keno -Magnathis. And you

00:13:11.610 --> 00:13:14.110
had Tenuacaptor. That completely changes the

00:13:14.110 --> 00:13:16.210
picture of that ancient world. Yeah. It wasn't

00:13:16.210 --> 00:13:18.909
a monoculture. It was a diverse community. Exactly.

00:13:18.909 --> 00:13:21.110
It's kind of like looking at a blurry photo of

00:13:21.110 --> 00:13:22.980
a crowd and thinking, everyone is wearing the

00:13:22.980 --> 00:13:25.480
exact same gray suit. Then you finally get the

00:13:25.480 --> 00:13:27.659
high res version and you realize, oh, that guy's

00:13:27.659 --> 00:13:29.519
wearing a vest, that guy has a hat, and that

00:13:29.519 --> 00:13:31.580
one is wearing a tuxedo. And that implies they

00:13:31.580 --> 00:13:33.919
were living differently, right? If they are distinct

00:13:33.919 --> 00:13:36.259
species, they probably aren't all eating the

00:13:36.259 --> 00:13:38.419
exact same lunch at the same time. That's the

00:13:38.419 --> 00:13:41.090
major ecological implication, yes. It strongly

00:13:41.090 --> 00:13:43.690
suggests niche partitioning. Niche partitioning.

00:13:43.710 --> 00:13:45.830
Right, meaning they divided up the resources

00:13:45.830 --> 00:13:49.269
so they didn't directly compete. Maybe Caenomagnathus

00:13:49.269 --> 00:13:51.429
has stronger jaws for cracking harder shells

00:13:51.429 --> 00:13:54.409
or taking down tougher prey. Maybe Tenuacaptor

00:13:54.409 --> 00:13:56.750
was lighter and faster and chased down quick

00:13:56.750 --> 00:14:00.049
insects. It shows us that this ecosystem 300

00:14:00.049 --> 00:14:02.809
million years ago was incredibly complex. It

00:14:02.809 --> 00:14:05.029
was a mature food web with different synapsid

00:14:05.029 --> 00:14:07.389
predators finding their own unique ways to survive.

00:14:07.690 --> 00:14:10.799
And all of this came from the exact pile of bones

00:14:10.799 --> 00:14:12.960
that have been sitting in museum drawers since

00:14:12.960 --> 00:14:15.759
the 1970s. That's the beauty of paleontology.

00:14:15.919 --> 00:14:18.200
The bones didn't change at all. Our questions

00:14:18.200 --> 00:14:20.519
changed. Our ability to see the fine details

00:14:20.519 --> 00:14:23.220
changed. So let's zoom out a bit and look at

00:14:23.220 --> 00:14:25.240
the evolutionary neighborhood. We've met the

00:14:25.240 --> 00:14:28.259
cast, Eoheptides, Keno Magnathus, Tenuacaptor.

00:14:28.480 --> 00:14:30.940
Where do they fit in the grand scheme? The source

00:14:30.940 --> 00:14:33.480
provides a cladogram, which is basically an evolutionary

00:14:33.480 --> 00:14:35.860
family tree, and it places them in a specific

00:14:35.860 --> 00:14:39.120
group called Sphynacodontia. Right. So Sphynacodontia

00:14:39.120 --> 00:14:41.659
is the club. It includes our Kansas trio. It

00:14:41.659 --> 00:14:43.539
includes the famous Dimetrodon we talked about

00:14:43.539 --> 00:14:45.779
earlier. It also includes other basal members

00:14:45.779 --> 00:14:49.240
mentioned in the source, like Cutleria. And looking

00:14:49.240 --> 00:14:51.139
at that tree, I see some other great names nearby.

00:14:51.600 --> 00:14:54.620
We have the clade Metapophora and then a group

00:14:54.620 --> 00:14:57.279
called Paleohatteridae. Yes. Paleohatteridae

00:14:57.279 --> 00:14:59.960
is a closely related family on the tree. It includes

00:14:59.960 --> 00:15:02.679
animals like Paleohatteria and the delightfully

00:15:02.679 --> 00:15:05.399
named Pantelosaurus. Pantelosaurus. Yeah. That

00:15:05.399 --> 00:15:07.240
sounds like a dinosaur that plays charades. It

00:15:07.240 --> 00:15:10.179
really does. Yeah. But these groups, Metapophora,

00:15:10.340 --> 00:15:14.580
Paleohatteridae, Sphenicodontia, they represent

00:15:14.580 --> 00:15:17.059
the critical steps on the evolutionary ladder.

00:15:17.200 --> 00:15:19.860
The source lays out the trajectory very clearly.

00:15:20.320 --> 00:15:23.100
You start broad with synapsida. That narrows

00:15:23.100 --> 00:15:25.220
down over millions of years into sphenicodontia.

00:15:25.299 --> 00:15:28.039
That evolves into therapsida. Then synodontia.

00:15:28.139 --> 00:15:31.960
And finally, mammals. Mammals. Us. These creatures

00:15:31.960 --> 00:15:34.860
scrambling around in the Kansas mud, chino magnathus,

00:15:34.860 --> 00:15:37.539
and eohaptidus, they were the experimental phase.

00:15:37.679 --> 00:15:39.860
The rough drafts. Exactly. The rough drafts of

00:15:39.860 --> 00:15:41.840
the mammalian blueprint. They were figuring out

00:15:41.840 --> 00:15:44.159
how to walk efficiently on land, how to process

00:15:44.159 --> 00:15:46.799
food better, how to begin regulating their bodies.

00:15:47.370 --> 00:15:49.850
It's humbling, honestly. You look at the skeletal

00:15:49.850 --> 00:15:51.990
reconstruction of something like Kena Magnathus,

00:15:52.029 --> 00:15:55.529
it has low slung sprawling legs, a big heavy

00:15:55.529 --> 00:15:58.529
head, a long tail, and it looks like a monster.

00:15:59.289 --> 00:16:02.210
But That's family. That's the great grand ancestors

00:16:02.210 --> 00:16:04.549
struggling in the mud so that we could eventually

00:16:04.549 --> 00:16:07.129
sit here and analyze their bones. And that is

00:16:07.129 --> 00:16:09.490
exactly why the taxonomy matters so much. That's

00:16:09.490 --> 00:16:12.049
why Spindler's incredibly tedious work to split

00:16:12.049 --> 00:16:14.549
them up is so important. If we just lump them

00:16:14.549 --> 00:16:16.970
all together as Heptotis, we lose the resolution

00:16:16.970 --> 00:16:20.389
of that history. We miss the diversity. By recognizing

00:16:20.389 --> 00:16:22.210
that there were three different species trying

00:16:22.210 --> 00:16:24.970
three different survival strategies, we literally

00:16:24.970 --> 00:16:27.370
see evolution in action. We are giving them their

00:16:27.370 --> 00:16:30.519
dignity back. even if their real names are currently

00:16:30.519 --> 00:16:32.840
stuck in a dissertation. Science is a process,

00:16:32.919 --> 00:16:35.700
not a destination. So let's recap this journey,

00:16:35.799 --> 00:16:37.419
because we've covered a lot of ground today and

00:16:37.419 --> 00:16:39.720
a lot of mud. We started with a simple question

00:16:39.720 --> 00:16:42.320
about a name in quotation marks. We traveled

00:16:42.320 --> 00:16:45.299
back 304 million years to the Garnett Quarry

00:16:45.299 --> 00:16:47.899
in Kansas. Where we found a nursery of fossils

00:16:47.899 --> 00:16:50.399
that we confidently thought were all one thing.

00:16:51.000 --> 00:16:54.000
Then we watched as modern science basically took

00:16:54.000 --> 00:16:57.240
a sledgehammer to that assumption. We saw the

00:16:57.240 --> 00:16:59.659
original Heptotis named Crumble because it was

00:16:59.659 --> 00:17:02.519
based on bad data from Europe. We saw Eoheptotis

00:17:02.519 --> 00:17:05.420
proposed as a replacement. And then the massive

00:17:05.420 --> 00:17:08.319
plot twist. We saw the population split again

00:17:08.319 --> 00:17:11.799
into Kinomagnathus and Tenuacaptor. We learned

00:17:11.799 --> 00:17:14.220
that a name on a plaque isn't a hard fact. It's

00:17:14.220 --> 00:17:16.200
just the best hypothesis we have at the time.

00:17:16.299 --> 00:17:19.519
That is the ultimate key takeaway here. Heptotis

00:17:19.519 --> 00:17:23.599
was a hypothesis that stood for 40 years. Eoheptotis

00:17:23.599 --> 00:17:26.519
and Kinomagnathus are the new hypotheses. They

00:17:26.519 --> 00:17:28.920
might hold up forever or, you know, in 50 years,

00:17:28.960 --> 00:17:30.680
someone with a new particle scanner might look

00:17:30.680 --> 00:17:32.839
at those exact same bones and find something

00:17:32.839 --> 00:17:35.299
spindler missed. It really makes you wonder what

00:17:35.299 --> 00:17:37.279
else is hiding in plain sight. That is the big

00:17:37.279 --> 00:17:39.180
question that keeps paleontologists awake at

00:17:39.180 --> 00:17:40.779
night. I mean, think about it for a second. We

00:17:40.779 --> 00:17:42.920
talked about the garnet quarry material, but

00:17:42.920 --> 00:17:44.500
think about the Smithsonian. Think about the

00:17:44.500 --> 00:17:46.599
Natural History Museum in London. The endless

00:17:46.599 --> 00:17:48.839
rows of cabinets in the basements that visitors

00:17:48.839 --> 00:17:52.059
never even get to see. There are millions of

00:17:52.059 --> 00:17:54.440
specimens in those collections. Millions. And

00:17:54.440 --> 00:17:57.440
many of them haven't been seriously, rigorously

00:17:57.440 --> 00:18:00.319
examined since the Victorian era. They were dug

00:18:00.319 --> 00:18:03.380
up, tagged, thrown in a drawer, and essentially

00:18:03.380 --> 00:18:05.720
forgotten. How many of them are wearing the wrong

00:18:05.720 --> 00:18:08.700
name tag? How many Hepatitis -style situations

00:18:08.700 --> 00:18:11.400
are sitting there right now? It is almost a statistical

00:18:11.400 --> 00:18:14.059
certainty that there are entirely new species,

00:18:14.359 --> 00:18:17.259
completely unknown animals, sitting in museum

00:18:17.259 --> 00:18:19.940
drawers right this second. They have already

00:18:19.940 --> 00:18:21.880
been discovered, they just haven't been identified

00:18:21.880 --> 00:18:25.299
yet. And that is a wild thought to end on. The

00:18:25.299 --> 00:18:27.500
next massive breakthrough in understanding our

00:18:27.500 --> 00:18:30.240
own origins might not come from a multi -million

00:18:30.240 --> 00:18:32.759
dollar expedition to the Gobi Desert. It might

00:18:32.759 --> 00:18:35.180
literally come from a grad student blowing the

00:18:35.180 --> 00:18:37.559
dust off a cardboard box in a basement in New

00:18:37.559 --> 00:18:40.079
York. The discovery is in the data. You just

00:18:40.079 --> 00:18:42.359
have to look close enough. Well, on that note,

00:18:42.380 --> 00:18:44.039
I think I'm going to look a lot more closely

00:18:44.039 --> 00:18:46.039
at those little brass labels the next time I'm

00:18:46.039 --> 00:18:48.660
at a museum. We are going to wrap up this deep

00:18:48.660 --> 00:18:51.380
dive into the messy, bureaucratic, and completely

00:18:51.380 --> 00:18:54.700
fascinating world of synapsid taxonomy. Thank

00:18:54.700 --> 00:18:56.619
you for helping us decode the quotation marks

00:18:56.619 --> 00:18:59.059
today. Always a pleasure. It's a good reminder

00:18:59.059 --> 00:19:01.680
to always check your sources. Stay curious, everyone.

00:19:02.460 --> 00:19:04.000
We'll catch you on the next deep dive.
